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    Biosynthesis, production site, and emission rates of aggregation-attachment pheromone in males of two Amblyomma ticks
    The aggregation-attachment pheromone componentso-nitrophenol (ONP) and methyl salicylate (MS) in male Amblyomma variegatum ticks appeared after three days of feeding on the host and reached high values after about six days. Variable quantities of 1.3ā€“7.3 Ī¼g ONP and about 0.6 Ī¼g MS were present within ticks. ONP and MS were released at the high rates of 300ā€“1800 ng/hr and 20ā€“600 ng/hr per male tick, respectively. After a temporary decrease, males continued to emit at high rates after nearby attachment of females. In A. hebraeum, ONP showed a similar pattern, but with a delay of about a day. A male, which had fed during 14 days, contained about 2 Ī¼g and released 225ā€“280 ng/hr. Emission in forcibly detached males of both species dropped rapidly to low levels of less than 10 ng/hr per tick. Host skin and tick feces in the vicinity of feeding males were pheromoneimpregnated. The very high emission rates are consistent with the observations that the pheromone is an important component of the host-location mechanism of conspecifics. ONP and MS are produced in the dermal glands type 2 associated with the ventrolateral cuticle.