<i>DE MACROCARPAEAE GRISEBACH (EX GENTIANACEIS) SPECIEBUS NOVIS </i>: IV: ELEVEN NEW SPECIES OF <i>MACROCARPAEA</i> (GENTIANACEAE: HELIEAE) FROM CENTRAL AND SOUTH AMERICA, AND THE FIRST REPORT OF THE PRESENCE OF STIPULES IN THE FAMILY

Progress toward a taxonomic revision of <i>Macrocarpaea</i> (Gentianaceae: Helieae) has revealed numerous novelties in this morphologically diverse and unexpectedly species-rich genus. Eleven new species are here described and illustrated from Central and South America: Colombia (<i>M. nicotianifolia</i> and <i>M. schultesii</i>), Costa Rica (<i>M. auriculata</i>), Ecuador (<i>M. subsessilis</i>), Peru (<i>M. gracilis</i>, <i>M. maguirei</i>, <i>M. wurdackii</i>, and <i>M. zophoflora</i>), and Venezuela (<i>M. ewaniana</i>, <i>M. papillosa</i>, and <i>M. weaveri</i>). <i>Macrocarpaea zophoflora</i> is notable for being the first species to be described of Gentianaceae that is known, or recognized, to have free deciduous interpetiolar stipules. The family was previously defined as exstipulate without exception.